#NEXUS [written Thu Nov 19 13:30:36 GMT 2015 by Mesquite version 3.04 (build 725) at ESCI-5BV0H62/157.140.110.4 (Greg Edgecombe)] BEGIN TAXA; TITLE Taxa; DIMENSIONS NTAX=55; TAXLABELS Shankouia_zhenghei Aglaspis_spinifer Agnostus_pisiformis Alalcomenaeus_cambricus Arthroaspis_bergströmi Australimicola_spriggi Burgessia_bella Campanamuta_mantonae Cheloniellon_calmani Cindarella_eucalla Emeraldella_brocki Emucaris_fava Eoredlichia_intermedia Eurypterida Flobertia_kochi Fortiforceps_foliosa Haikoucaris_ercaiensis Helmetia_expansa Henningsmoenicaris_scutula Isoxys_volucris Jianfengia_multisegmentalis Kangacaris_zhangi Kiisortoqia_soperi Kuamaia_lata Kwanyinaspis_maotianshanensis Leanchoilia Liwia Marrella_splendens Martinssonia_elongata Mimetaster_hexagonalis Misszhouia_longicaudata Molaria_spinifera Naraoia_compacta Naraoia_spinosa Nebalia Oelandocaris_oelandica Olenoides_serratus Phytophilaspis_pergamena Pycnogonida Rebachiella_kinnekullensis Retifacies_abnormalis Saperion_glumaceum Sidneyia_inexpectans Sinoburius_lunaris Siriocaris_trollae Skioldia_aldna Soomaspis_splendida Squamacula_clypeata Tariccoia_arrusensis Tegopelte_gigas Triarthrus_eatoni Weinbergina_opitzi Xandarella_spectaculum Yohoia_tenuis Eozetetes_gemmelli ; END; BEGIN CHARACTERS; TITLE Character_Matrix; DIMENSIONS NCHAR=74; FORMAT DATATYPE = STANDARD GAP = - MISSING = ? SYMBOLS = " 0 1 2 3 4"; CHARSTATELABELS 1 'Nature of first cephalic appendage: (0) limb-like appendage; (1) antennae.', 2 'Composition of first cephalic appendage: (0) 7?15 articles; (1) > 15 articles; (2) ? 7 articles.', 3 'Armature of first cephalic appendage: (0) none; (1) pair of simple, robust spines medioventrally on each article; (2) setae on mediodistal margin of articles; (3) single, large, medodistal spine or finger.', 4 'Number of articles with fingers in short feeding appendage: (0) four; (1) three; (2) two.', 5 'Long spinose projections on distal part of terminal three podomeres of great appendage bearing a flagellum: (0) absent; (1) present.', 6 'Feeding appendage with elbow joint: (0) absent; (1) present.', 7 'Number of appendage bearing post-ocular segments incorporated into the head/prosoma: (0) one; (1) four; (2) five; (3) six; (4) seven. ', 8 'Exopod on first post-antennular appendage: (0) present; (1) absent.', 9 'Exopod on second post-antennular appendage: (0) present; (1) absent.', 10 'Insertion of cephalic exopods: (0) as in trunk; (1) shorter joint.', 11 'Cephalic endopods: (0) as in trunk limbs; (1) first postantennular limb with fewer podomeres or reduced; (2) endopod of first postantennular limb heavily reduced or absent, of second postantennular limb considerably more slender than in trunk and fewer podomeres.', 12 'Composition of first and second cephalic exopods: (0) as in trunk limbs; (1) multiarticulate, each article with mediodistal setae or pair of lateral setae.', 13 'Trunk endopods: (0) absent or reduced; (1) present.', 14 'Exopod structure: (0) simple oval flap; (1) bilobate flap, exopod differentiated into proximal and distal lobes; (2) numerous articles; (3) book gills.', 15 'Proximal lobe of exopod: (0) flattened lobe; (1) slender shaft.', 16 'Distal lobe of exopod: (0) small to moderate sized flap; (1) large, teardrop shaped lobe with long attachment; (2) dominant part of appendage, shielding endopod.', 17 'Joints in flap-like exopod (0) absent; (1) one; (2) two.', 18 'Imbricate exopod lamellae: (0) absent; (1) present.', 19 'Non-overlapping marginal setae: (0) absent; (1) small setae; (2) long spines.', 20 'Insertion of exopod on trunk appendages: (0) along lateral edge of multiple podomeres of limb axis; (1) on basipod and first endopod podomere; (2) on basipod only.', 21 'Setae of anterior exopods facing endopods: (0) absent; (1) present.', 22 'Body-limb joint: (0) short, sclerotized, pivot-jointed rings; (1) arthrodial membrane with partially sclerotized half-rings; (2) arthrodial membrane.', 23 'Endites on basipod/gnathobase: (0) absent; (1) multiple spinose endites; (2) single spinose endite.', 24 'Composition of endopod: (0) nine or eight podomeres; (1) seven podomeres; (2) fewer than seven podomeres.', 25 'Median armature of podomeres: (0) none; (1) mediodistal spines or spinules/denticles on all but penultimate and distal podomeres; (2) biserial spines or spinose endites along median edge of podomeres 1?4.', 26 'Proximal endite or coxa as separate sclerotized element proximal to limb base in cephalic appendages (0) absent; (1) present.', 27 'Position of lateral eyes: (0) ventral; (1) dorsal.', 28 'Visual surface with calcified lenses, bounded with circumocular suture: (0) absent; (1) present.', 29 'Dorsal bulge in exoskeleton accommodating drop-shaped ventral eyes: (0) absent; (1) present.', 30 'Eye slits: (0) absent; (1) present.', 31 'Dorsal median eyes: (0) absent; (1) present.', 32 'Bilobate lateral eyes where four visual surfaces are arranged in a subtransverse band across head shield: (0) absent; (1) present.', 33 'Expanded cephalic doublure: (0) absent; (1) concentric; (2) crescentic.', 34 'Anteromedian margin of cephalon notched, accommodating strongly sclerotised plate: (0) absent; (1) present.', 35 'Hypostomal sclerite: (0) wide attachment with or without suture; (1) natant; (2) with narrow overlap with prehypostomal sclerite; (3) absent.', 36 'Hypostome divided into anterior and posterior parts by transverse suture: (0) absent; (1) present.', 37 'Labrum: (0) absent; (1) present.', 38 'Prehypostomal sclerite: (0) as individual sclerite anterior to head tergite or recessed in notch in head shield; (1) incorporated in doublure as rostral plate or reduced.', 39 'Frontal organs on prehypostomal sclerite: (0) absent; (1) present.', 40 'Visible ecdysial dorsal sutures: (0) absent; (1) present.', 41 'Position of ecdysial sutures: (0) marginal; (1) dorsal.', 42 'Elevated marginal rim: (0) absent; (1) present.', 43 'Distinct trilobation of head shield, marked by axial furrows: (0) absent; (1) present.', 44 'Dorsal expression of last segment of head shield: (0) absent; (1) present.', 45 'Head shield outline: (0) genal angles, rounded, acute or with spines; (1) lateral spine-like extensions of the head shield; (2) lateral margin extending into genal spines flanking anterior trunk.', 46 'Mineralized cuticle: (0) absent; (1) calcitic; (2) phosphatic.', 47 'Trunk tergites with expanded tergopleurae covering appendages dorsally: (0) absent; (1) present.', 48 'Free thoracic tergites: (0) absent; (1) present.', 49 'Decoupling of thoracic tergites and segments: (0) absent; (1) present.', 50 'Tergite articulations: (0) tergites non-overlapping; (1) extensive overlap of tergites; (2) edge-to-edge pleural articulations.', 51 'Cephalic articulation fused: (0) absent; (1) present.', 52 'Head shield overlap of thoracic tergites: (0) overlap absent or identical to overlap between thoracic segments; (1) head shield covers first thoracic tergite only; (2) head shield covers multiple anterior trunk tergites.', 53 'Trunk narrowed anteriorly relative to head shield, widest posteriorly: (0) absent; (1) present.', 54 'Raised axial region of trunk defined by axial furrows: (0) absent; (1) present.', 55 'Articulating device: (0) articulating half-ring and flanges; (1) articulating ridge/anterior process; (2) no skeletal device.', 56 'Postcephalic tagmosis: (0) uniform trunk with free tergites and telson or pygidium; (1) bipartite trunk with free tergites and telson, segments in abdomen apodous, tergites ring-shaped, without tergopleurae.', 57 'Trunk segments form single shield: (0) absent; (1) present.', 58 'Posterior tergites strongly curved in dorsal aspect compared to anterior tergites: (0) absent; (1) present.', 59 'Pygidium: (0) absent; (1) present.', 60 'Position of the anus: (0) terminal; (1) ventral.', 61 'Pygidium with median keel: (0) absent; (1) present.', 62 'Pygidium with broad-based median spine: (0) absent; (1) present.', 63 'Pygidium with lateral spines: (0) absent; (1) present.', 64 'Size of caudal end: (0) shorter to slightly longer than cephalon; (1) much longer than cephalon.', 65 'Free telson: (0) present; (1) absent.', 66 'Telson shape: (0) styliform keeled; (1) plate-like; (2) tubular cap-like to tubular styliform.', 67 'Telson carrying furca: (0) absent; (1) present.', 68 'Spines on telson: (0) absent; (1) present.', 69 'Segmentation of telson: (0) absent (unjointed); (1) present.', 70 'Postventral plates: (0) absent; (1) present.', 71 'Paired modified appendages attached to a differentiated pre-terminal segment: (0) absent; (1) present.', 72 'Nature of pre-terminal appendages: (0) paddle; (1) cerci.', 73 'Relative length of thorax: (0) shorter than caudal end; (1) longer than caudal end.', 74 'Differentiation of cephalic gut diverticulae: (0) small, serially regular diverticulae; (1) anterior pair of diverticulae large, expanding across genal region of cephalic shield, with complicated ramification.' ; MATRIX Shankouia_zhenghei 000???0000?010??001000??000000?010???000?0000011010200?00001???0010000??0? Aglaspis_spinifer ??????1?????1?????????????10000010000100?1002211010000100001???00200010?0? Agnostus_pisiformis ?02???100121110010210221200000000010010001?1011102001?000?1100?01????00?1? Alalcomenaeus_cambricus 02311110000010??102101101000000100100100?0000011010000?00001???00101000?00 Arthroaspis_bergströmi 112???1?0??011011111012120000000001001?0?0110011020000000?1100101????0??00 Australimicola_spriggi 1?????????????????????????00000??0?0?1?0?0000011010000?001?????0?????0??0? Burgessia_bella 112???100???1??????????12?0??00??0???1?0?00000010?0200?00?01???00200100?00 Campanamuta_mantonae 11????3?????1??????????????00000?0?????0?0000011020000000?1100101????0??00 Cheloniellon_calmani 11????111???11????1?0?21?0?00000001?01?0?00000110100011?010????0020000110? Cindarella_eucalla 112???2000001100111?0??1??00000000100100?0000011110200??00?1???0?????00?00 Emeraldella_brocki 112???1?0010110021110121200??00000000?00?0000011010000100001???00200101000 Emucaris_fava 1?????1?????1?????????????000000001101?0?0000011??0001?00?1100011????0??1? Eoredlichia_intermedia 112???1000?0110001120?2120110000100001011111211102000100001100001????00?00 Eurypterida 023201311???03???0???220001000101030010101000011010000110001???00000000?0? Flobertia_kochi ????????????1?????????????10000010000100?10?2211?100001?0?0????0????010?0? Fortiforceps_foliosa 0230011000?010???0110?????00000?00?????0?0000011010000?00001???0010?000?0? Haikoucaris_ercaiensis 023101100??010???0??0?????00000??0???1?0?0000011010000?00001???00101000?0? Helmetia_expansa ??????1????0????????0?????00?00011200000?0000011020000000?1101101????00?0? Henningsmoenicaris_scutula 002???20012110??1021122?2100000000100100?0000011010000200??1???0?2?0000?0? Isoxys_volucris 001?????????10???01???????00000010???1?0?10000??????0??00?01????0????0??0? Jianfengia_multisegmentalis 023001100??010???01?0?????000000?0?????0?0000011010000?00001???00?0?000?0? Kangacaris_zhangi 1?????1?0???1?????????????0000?0001101?0?0000011?20000?00?1100011????0??1? Kiisortoqia_soperi 001??0100?0010???01?01?0?000000??0???1?0?00000110100001000?1???0?????00?00 Kuamaia_lata 112????00???110111?1012120001000112000?0?0000011020000000?1101101????00?0? Kwanyinaspis_maotianshanensis ???????00???110111110??120001000000??1?0?00?0011020000000001???002000?0?0? Leanchoilia 02311110000010??1021011010000001001?0100?0000011010000100001???00101000?00 Liwia 11????????????????????????????0??0???1???00000110?0110000?1110111????00?1? Marrella_splendens 112????00?1012???1?????12????00?00???1?0?00010010?0?00?00?0????0??0??00?0? Martinssonia_elongata 022??020000110???022122221?0000000?00100?0000001000000210?00???0?21?000?0? Mimetaster_hexagonalis 11?????????012?????????12?00000?001??1?0?00010010?0?00?00?0????0??0??00?0? Misszhouia_longicaudata 112???2000001110111101212000000000100110?00000100?0010?01?1100011????00?10 Molaria_spinifera 102???1?????1100?11????12?00000000???1?0?0000011010000100001???0020010??0? Naraoia_compacta 112????000?01110111?0?212?00000000100110?00000100?0010?01?1100011????00?11 Naraoia_spinosa 112???2000?01101011101212000000000100110?00000100?0010?01?1100111????00?11 Nebalia 112???200?0010???012?2222100000000101100?0000001000200210?00???0?21?000?00 Oelandocaris_oelandica 0?????20012110??1021122?2100000000100100?0000001000000200??1???0?200000?0? Olenoides_serratus 112???10000011001112012120110000100001011111211102000100001100101????0110? Phytophilaspis_pergamena ??????2???????????????????10010??0?0?1?0?0?12211??0001000?1100011???????1? Pycnogonida 023201311????????0???20010?00010003?0100?0?0?0???????????????????????00??0 Rebachiella_kinnekullensis 022??020002110??001212122100000000101100?0000001000200210?00???0?21?000?0? Retifacies_abnormalis 112???100??01???01?101?12?000000?0000?00?000001101000010001?0???0200100?0? Saperion_glumaceum 1???????????110111?101?1??001000112000?0?00000100?1000000?1100001????00?0? Sidneyia_inexpectans 112?????????11022111??212?0000001000010??0000011010000110001???0010000100? Sinoburius_lunaris ????????????1?????????????00??0000???1?0?01?20110?0000?00?1101101????00?0? Siriocaris_trollae 112???100??0110111?10?2?2?000000000001?0?0000011020000000001???0010000??00 Skioldia_aldna 112?????????1????1???1????0010001120?0?0?0000011?20000000?1100001????00?0? Soomaspis_splendida ??????????????????????????0?000?20???1?0?0000011?20110000?1110111????00?1? Squamacula_clypeata 11??????????1??2??1???21??000000?000?100?0000011010000??00?1???0?????00??? Tariccoia_arrusensis ??????????????????????????0?000?20???1?0?0000011?20110?00?1110111????00?1? Tegopelte_gigas 112????000??1100?1??0??1??0??000?1?0???0?000?010??1?00?00?11000?1????00??? Triarthrus_eatoni 11????1000001????1??0?2120110000100001011111211102000100001100001????0??0? Weinbergina_opitzi 023201311???03???01??22?001000101030010101000011010000110001???00000000?0? Xandarella_spectaculum 112???4000??1110111?0?2?2000010000100100?0000011110200??00?1???0?????00?00 Yohoia_tenuis 0230011000?010???01?0?10??00000000??0??0?0000011010000?10001???00101000?0? Eozetetes_gemmelli 11????????????????????????0?0000100001?0?1000011?10000?00001???0020000??0? ; END; BEGIN ASSUMPTIONS; TYPESET * UNTITLED = unord: 1 - 74; END; BEGIN MESQUITECHARMODELS; ProbModelSet * UNTITLED = 'Mk1 (est.)': 1 - 74; END; Begin MESQUITE; MESQUITESCRIPTVERSION 2; TITLE AUTO; tell ProjectCoordinator; timeSaved 1447939836298; getEmployee #mesquite.minimal.ManageTaxa.ManageTaxa; tell It; setID 0 293120180604408739; endTell; getEmployee #mesquite.charMatrices.ManageCharacters.ManageCharacters; tell It; setID 0 365761748539148046; mqVersion 304; checksumv 0 3 2672561714 null getNumChars 74 numChars 74 getNumTaxa 55 numTaxa 55 short true bits 31 states 31 sumSquaresStatesOnly 9032.0 sumSquares 9032.0 longCompressibleToShort false usingShortMatrix true NumFiles 1 NumMatrices 1; mqVersion; endTell; getWindow; tell It; suppress; setResourcesState false false 100; setPopoutState 400; setExplanationSize 0; setAnnotationSize 0; setFontIncAnnot 0; setFontIncExp 0; setSize 1284 630; setLocation -8 0; setFont SanSerif; setFontSize 10; getToolPalette; tell It; endTell; desuppress; endTell; getEmployee #mesquite.minimal.ManageTaxa.ManageTaxa; tell It; showTaxa #293120180604408739 #mesquite.lists.TaxonList.TaxonList; tell It; setTaxa #293120180604408739; getWindow; tell It; newAssistant #mesquite.lists.TaxonListCurrPartition.TaxonListCurrPartition; newAssistant #mesquite.lists.DefaultTaxaOrder.DefaultTaxaOrder; getTable; tell It; rowNamesWidth 223; endTell; setExplanationSize 30; setAnnotationSize 20; setFontIncAnnot 0; setFontIncExp 0; setSize 1184 558; setLocation -8 0; setFont SanSerif; setFontSize 10; getToolPalette; tell It; setTool mesquite.lists.TaxonList.TaxonListWindow.ibeam; endTell; setActive; endTell; showWindow; getEmployee #mesquite.lists.ColorTaxon.ColorTaxon; tell It; setColor Red; removeColor off; endTell; getEmployee #mesquite.lists.TaxonListAnnotPanel.TaxonListAnnotPanel; tell It; togglePanel off; endTell; endTell; endTell; getEmployee #mesquite.charMatrices.BasicDataWindowCoord.BasicDataWindowCoord; tell It; showDataWindow #365761748539148046 #mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindowMaker; tell It; getWindow; tell It; getTable; tell It; rowNamesWidth 208; endTell; setExplanationSize 30; setAnnotationSize 20; setFontIncAnnot 0; setFontIncExp 0; setSize 1184 558; setLocation -8 0; setFont SanSerif; setFontSize 10; getToolPalette; tell It; setTool mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindow.ibeam; endTell; setTool mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindow.ibeam; colorCells #mesquite.charMatrices.NoColor.NoColor; colorRowNames #mesquite.charMatrices.TaxonGroupColor.TaxonGroupColor; colorColumnNames #mesquite.charMatrices.CharGroupColor.CharGroupColor; colorText #mesquite.charMatrices.NoColor.NoColor; setBackground White; toggleShowNames on; toggleShowTaxonNames on; toggleTight off; toggleThinRows off; toggleShowChanges on; toggleSeparateLines off; toggleShowStates on; toggleAutoWCharNames on; toggleAutoTaxonNames off; toggleShowDefaultCharNames off; toggleConstrainCW on; toggleBirdsEye off; toggleShowPaleGrid off; toggleShowPaleCellColors off; toggleShowPaleExcluded off; togglePaleInapplicable on; toggleShowBoldCellText off; toggleAllowAutosize on; toggleColorsPanel off; toggleDiagonal on; setDiagonalHeight 80; toggleLinkedScrolling on; toggleScrollLinkedTables off; endTell; showWindow; getWindow; tell It; forceAutosize; endTell; getEmployee #mesquite.charMatrices.ColorByState.ColorByState; tell It; setStateLimit 9; toggleUniformMaximum on; endTell; getEmployee #mesquite.charMatrices.ColorCells.ColorCells; tell It; setColor Red; removeColor off; endTell; getEmployee #mesquite.categ.StateNamesStrip.StateNamesStrip; tell It; showStrip off; endTell; getEmployee #mesquite.charMatrices.AnnotPanel.AnnotPanel; tell It; togglePanel off; endTell; getEmployee #mesquite.charMatrices.CharReferenceStrip.CharReferenceStrip; tell It; showStrip off; endTell; getEmployee #mesquite.charMatrices.QuickKeySelector.QuickKeySelector; tell It; autotabOff; endTell; getEmployee #mesquite.charMatrices.SelSummaryStrip.SelSummaryStrip; tell It; showStrip off; endTell; getEmployee #mesquite.categ.SmallStateNamesEditor.SmallStateNamesEditor; tell It; panelOpen true; endTell; endTell; endTell; endTell; end;